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the pressure instantly equalizes and the syringe volume is displaced. Same with a forced induction engine. The displacement and rpm control the volume of air. There is a slight pressure drop across the intercooler, and thru the piping, and that causes a difference in the mass per volume at those locations. Of course different masses are made up of different volumes of intake air. The engine displaces about 200 cfm per side at redline. With the slight drop in pressure across the intercooler, the volume of air is slightly less before the intercooler, but this is more than offset when the air temp is reduced by heat transfer and the air contracts.
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